Photographs of Grand Teton National Park from summer 2011
Grizzly 399 retreats to the woodland edge to nurse her triplet cubs.
Grizzly 399 reacts to an overeager cub who has nursed a bit too roughly.
Tired of her cub's bites, grizzly 399 gives one of her triplet cubs an old fashioned smackdown in June 2011.
Girzzly 399 and her three cubs forage in a meadow of bitterroot and lupines.
Grizzly 399, the grand matriarch of the Grand Tetons, remains on high alert as one of her three cubs checks out the meadow's flowers in the summer of 2011.
Grizzly matriarch 399 leads her three young cubs across the meadow and into the forest where they will spend the majority of hte day nursing and napping.
399 nourishes her triplet cubs of the year.
399 has been a prolific and effective mom, in part because of her constant care and nourishment of her cubs over two decades.
Grizzly moms, like 399, know the importance of administering swift and firm lessons to wayward cubs.
A pond of water, the final remnants of winter's heavy snows, is slow to drain in a spring meadow, reflecting twilight's fading colors.
Grizzly 399 and one of her three cubs march slowly across a sage meadow.
Two of grizzly 399's three cubs play on fallen timbers.
A small stream cascades across the stones, just downstream from the beaver dam that defines Schwabacher's Landing.
Shortly after sunrise, the sun has now reached the still waters of a beaver pond in the shadows of the pines below the still snow-capped peaks of the Grand Tetons.
One of my most popular images for more than a decade, Moulton Barn is warmed by the light of an earl summer sunrise, with snow-capped mountains and still-green grasses in vibrant display.
Grizzly 399 and one of her three cubs enjoy a quiet morning in the sage and biterroot meadow.
A young 399 forages in the meadow as her three cubs play nearby.
Grizzly 399 approaches through the wildflowers.
Grizzly 399 looks into the nearby woods where she has detected some sound or smell.
Two of grizzly 399's cubs play on a meadow post.
A purple majesty at dawn at Oxbow Bend.
Lupines bloom amidst a field of bitteroot.
Snow-capped peaks of the Teton mountain range rise above the slopes of Signal Mountain.
Grizzly 610 stalks elk in an open meadow near Willow Flats. Not only are her two cubs anxious for success, but just out of view, a black wolf watches with equal interest. These are the moments that you never forget in the Grand Tetons.
A field of flowers along Antelope Flats lean toward the rising sun.
Dawn's light bathes Oxbow Bend and the Teton range in a sea of pastel blues and purples.
Dawn's first rays light the peak of Mount Moran and the Teton Range above Oxbow Bend.
Few places are more magical to watch sunrise than the beaver ponds at Schwabacher's Landing. Winter's deep snows provide a powerful backdrop even in late June.
Morning on the Oxbow Bend is gradually revealed as the sun strikes Mount Moran. Ducks, geese and assorted widlife cavort in and along the water's edge in one of the most wildlife-rich areas of the Grand Tetons.