She may be just over 20 years of age, but grizzly 399 remains an indomitable force.

Still Going Strong

Grand Teton National Park

She may be just over 20 years of age, but grizzly 399 remains an indomitable force. Photo © copyright by Ed Fuhr.

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Related Galleries: Bears, Springtime with 399 and Blondie in the Tetons 2018 , Grizzly 399 Photography Collection - Queen of the Tetons