Grizzly 399, now some 24 years old and the grand matriarch of the Grand Tetons, is undoubtedly the most famous grizzly bear in...

Matriarch

Grand Teton National Park

Grizzly 399, now some 24 years old and the grand matriarch of the Grand Tetons, is undoubtedly the most famous grizzly bear in the world, in large measure because she has seeded the Tetons with her offspring and raised them at times near the Park's roadways. Photo © copyright by Ed Fuhr.